Annuals; (usually glaucous distally), often sparsely to densely hirsute proximally, sometimes glabrous throughout. Stems unbranched or branched throughout, 1.5-12 dm. Basal leaves (soon withered); not rosulate; petiolate (petiole winged); blade lanceolate to oblanceolate, 1-15 cm, base narrowed to petiole, margins coarsely dentate to ± lobed. Cauline leaves: blade linear-lanceolate to lanceolate or oblanceolate, 1-12 cm × 1-11 mm, (smaller distally), blade auriculate to amplexicaul, margins entire or coarsely dentate, (often entire distally). Racemes ebracteate, (lax, secund or not, rachis usually straight, rarely flexuous). Fruiting pedicels ascending to divaricate or spreading, (straight or recurved), 2-32 mm, (glabrous or pubescent). Flowers (zygomorphic); calyx urceolate; sepals white, greenish white, cream, yellow, rose, purple, lilac, maroon, reddish purple, or purple-black, (lanceolate to broadly ovate), (3-)5-10 (-13) mm, not keeled, (glabrous or sparsely hirsute); petals lavender, purple, or white (sometimes with brown or purple veins), 7-17 mm, blade 2-7 × 1-3 mm, (subequal or adaxial pair distinctly longer, more recurved), margins crisped, claw 5-13 mm, wider than blade; stamens in 3 unequal pairs; filaments: abaxial pair (distinct), 4-9 mm, lateral pair 2.5-7 mm, adaxial pair (exserted, connate 2/3 their length), 5-13 mm; anthers: abaxial and lateral pairs fertile, 1-2.5 mm, adaxial pair sterile, 0.3-1 mm; gynophore 0.2-1.5 mm. Fruits ascending to divaricate or spreading, smooth, straight, curved upward, or arcuate, flattened, 3-11 cm × 1.5-2.5 mm; valves each with prominent midvein, (glabrous or sparsely to moderately hirsute); replum straight; ovules 22-70 per ovary; style 0.1-2.5 mm; stigma subentire. Seeds ovoid to oblong, 1.5-2.1 × 1-1.5 mm; wing 0.1-0.5 mm wide, continuous.