Perennials [shrubs], 10-60+ [-700] cm (rootstocks bearing tubers). Stems erect, branched ± throughout. Leaves cauline; opposite; petiolate; blades (usually 3-nerved) rounded-deltate to ovate or lance-ovate [lanceolate], bases cuneate to rounded or truncate, margins serrate, faces ± scabrous. Heads radiate, borne singly [in corymbiform arrays]. Involucres hemispheric [obconic], [4-]10-14[-25+] mm diam. Phyllaries persistent, 12-20 in 2-5 series (ovate to lanceolate, subequal or outer shorter [longer], herbaceous to papery). Receptacles convex, paleate (paleae membranous to indurate, conduplicate, apices acute [erose]). Ray florets [4-]11-14[30], pistillate, fertile; corollas yellow to orange [purplish]. Disc florets 20-100+[-200+], bisexual, fertile; corollas yellow to orange [purplish], tubes about equaling ampliate, cylindric throats, lobes 5, deltate. Cypselae 3-angled (ray) or strongly compressed or flattened (all with sharp edges, not truly winged); pappi persistent, of 2(-3) subulate scales or awns (continuous with margins of cypselae) plus 0-4 shorter scales. x = 10, 11-